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From July to August 2002, she performed on stage in Eve Ensler's The Vagina Monologues, She returned to Saturday Night Live on May 5, appearing as then-President George W.

Bush's daughter Jenna Bush in a skit that poked fun at the two first daughters for being arrested for underaged drinking.

Rolling Stone pronounced her "the coolest co-ed", putting her on the cover of its April 12, 2001, issue.

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She also had small roles as Harrison Ford's character's daughter in Alan J. Critic Joe Balthai wrote she was "the darling of the 1998 Sundance Film Festival." In 1999, she portrayed Kat Stratford, opposite Heath Ledger, in Gil Junger's 10 Things I Hate About You, an adaptation of The Taming of the Shrew set in a high school in Tacoma, Washington.

She won an MTV Movie Award for Breakthrough Female Performance for the role.

She had a recurring role as Lumen Pierce in the fifth season of the Showtime series Dexter (2010), a role that earned her nominations for the Golden Globe Award for Best Supporting Actress – Series, Miniseries or Television Film and the Primetime Emmy Award for Outstanding Guest Actress in a Drama Series.

As a member of the cast of Silver Linings Playbook (2012), Stiles was nominated for a Screen Actors Guild Award for Outstanding Performance by a Cast in a Motion Picture.

Stiles appears in the Blue web series also broadcast on television (2012–present).

Other film roles include romantic comedy The Prince and Me (2004), noir-inspired drama-thriller A Little Trip to Heaven (2005), British-Canadian co-production The Cry of the Owl (2009), Between Us (2012), Out of the Dark (2014), and with Anthony Hopkins in Blackway (2016) (also known as Go With Me). Her first lead was in Wicked (1998), playing a teenage girl who might have murdered her mother so she could have her father all to herself.

She has an effect on people." Stiles also had a small but crucial role as Treadstone operative Nicolette "Nicky" Parsons in The Bourne Identity (2002), a role that was enlarged in The Bourne Supremacy (2004), then greatly expanded in The Bourne Ultimatum (2007).

Between the Bourne films, she appeared in Mona Lisa Smile (2003) as Joan, a student at Wellesley College in 1953, whose art professor (Julia Roberts) encourages her to pursue a career in law rather than become a wife and mother.

Her first film role was in I Love You, I Love You Not (1996), followed by a leading role in the thriller Wicked (1998), for which she was awarded the Karlovy Vary International Film Festival Award for Best Actress.

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